Remember just because you have a reservation, it does not mean there is an actual car waiting. If you arrive at night, many of the cars may be gone and you may get a different car (upgrade and sometimes a downgrade). However, they will still honor your price.
I would suggest calling the rental agency on day of travel (if possible) and just let them know you were delayed and will be arriving at a later time. This way they can have a vehicle waiting for you if they have it (or a suitable upgrade). If they ask for flight numbers/times, tell them the info for the last leg (arrival into MCO/TPA). Flights get delayed all the time and people get shifted to different flights, as well. Standard activity at the airport, so they are used to it.